Actualllyyyyy I think i know what:
means. Its the information of what bone mirror is its equal. So bone NPC R Finger42 should be a mirror of NPC L Finger42 for example. The numbers represent what bone in the chain of the skeleton file is a mirror of the other.
I'm not sure this would do anything for you the only Mirrored Animations are the Walk/Run/Turn L/R's being mirrored that is why most actors only possess one or the other a Left or a Right, some have both -> this should not cause crashing -> for instance for my own Creatures I have been setting this array to 0 nodes and wiping out the array for testing my animations.
I do not think there is anything in the MetaData that needs altered its suppose to be nothing but hkClass data, also I have been deleting this out of my converted files with no ill omens haunting me -> with exception to my creatures not being resurrectable without disable/enable-ing them.
Oddly enough just the act of converting the female character profile to and fro breaks it.... not having this issue with the slaughterfish or werewolf
profiles which are the 2 profile sets I have converted for my custom creatures.
The more I fiddle with this the more I think it has something to do with your meshes, I just can't nail down whats wrong -> there are a few differences your ShapeData has normals and no VertexColors and the base has the exact opposite but even correcting these differences made no difference the game still crashed on those two races and not the others.